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- https://chemistry-europe.onlinelibrary.wiley.com/doi/abs/10.1002/1521-3765%2820011105%297%3A21%3C4553%3A%3AAID-CHEM4553%3E3.0.CO%3B2-0
- https://pubs.rsc.org/en/content/articlelanding/2004/pp/b316210a/unauth#!divAbstract
Oxidized alpha acids (humulinones) will skunk if exposed to UV light <ref>[https://www.homebrewersassociation.org/how-to-brew/resources/conference-seminars Dr. Patricia Aron. "Bitterness and the IBU: What’s It All About?" HomebrewCon 2017 Presentation. ~32 mins in. Retrieved 09/05/2017.]</ref>.