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====Thiols====
[https://researchmap.jp/?action=cv_download_main&upload_id=110987 Kishimoto et al.] found an increase in the thiol 3-methyl-2-butene-1-thiol (MBT) in beers that were brewed with unidentified pellet hops (described in the study only as "a bitter variety of 11.5% alpha acid") that were aged for 30 days at 40°C versus using fresh pellet hops that were stored cold (4°C). Interestingly, this thiol was higher in beers where the aged hops were added to the boil rather than when they were added after the wort was cooled. The authors were not able to determine whether or not the MBT was derived from yeast fermentation, or from boiling the hops, but aging the hops increased the precursors for MBT <ref name="kishimoto_2007" />. MBT has been described as the thiol that produces the "skunky" aroma in lightstruck beer <ref>[http://www.aroxa.com/beer/beer-flavour-standard/3-methyl-2-butene-1-thiol/ 3-methyl-2-butene-1-thiol. Aroxa website. Retrieved 11/22/2016.]</ref>.
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