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updated Sugar Utilization and Secondary Metabolites
===Sugar Utilization and Secondary Metabolites===
''Lactobacillus'' generally prefers glucose, fructose, and maltose, and does not ferment maltotriose. Some species may prefer certain types of sugars over others, for example ''L. plantarum'' ferments glucose first, and then fructose if it is available. ''L. reuteri'' ferments maltose first, while ''L. brevis'' feeds on maltose, glucose, and fructose. Peak sugar consumption without competition from yeast is typically 48 hours, and very little alcohol is produced (around 0.10-0.30%, far less than the 0.5% required for non-alcoholic drinks). Consumption of sugars occurs mainly during the 48 hour growth period, but also occurs after growth has stopped. No more than 0.5-1°P worth of sugar is consumed by ''Lactobacillus'', but rather than high residual sugar concentration being the limiting factor on growth it is thought that low pH and other metabolic byproducts weaken and finally stop the growth of ''Lactobacillus'' <ref name="Peyer"></ref>. For a chart and in depth discussion on what types of sugars are fermentable by different species of ''Lactobacillus'', as well as charts on ''secondary metabolites'', see [http://phdinbeer.com/2015/04/13/physiology-of-flavors-in-beer-lactobacillus-species/ Matt Humbard's ''Physiology of Flavors in Beer – Lactobacillus Species'' blog article]. Heterofermentative species can also produce [[Tetrahydropyridine]].
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